This Accepted Manuscript (published on 29 March 2022) is currently undergoing further peer-review before the final Version of Record is published.

After the acceptance of the manuscript, the authors requested to correct Figures 4, 5, 6 and 7. Given the significant corrections requested, the Accepted Manuscript and revised Figures are undergoing additional peer-review before the article's final publication. The Figures presented in the Accepted Manuscript may therefore be replaced in the Version of Record.

This is an open access article published by Portland Press Limited on behalf of the Biochemical Society and distributed under the Creative Commons Attribution License 4.0 (CC BY).